Suffering is a state of mind that is difficult to measure and analyse in human beings and considerably more so in animals. It is related to the environment in which we live and our physical and mental states. Understanding the physiology of suffering in animals is crucial in assessing animal welfare.

Written by an expert in applied welfare aspects of physiology, this book is the first to address the physiological aspects of suffering in animals. It explores the different causes of suffering – physical discomfort, thirst and hunger, the responses in the body that lead to suffering and it offers insight into how suffering can be managed.

The second book in a major new animal welfare series

Draws together information that is scattered across the literature

Written for the specialist and non-specialist alike

Includes colour pictures This book is part of the UFAW/Wiley-Blackwell Animal Welfare Book Series. This major series of books produced in collaboration between UFAW (The Universities Federation for Animal Welfare), and Wiley-Blackwell provides an authoritative source of information on worldwide developments, current thinking and best practice in the field of animal welfare science and technology. For details of all of the titles in the series see www.wiley.com/go/ufaw .

Professor Neville Gregory is based at the Royal Veterinary College, Hawkshead, London.